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The heads are still improved by the upgrades that affect all GP improvements in general (i.e. freedom tenant, new deal policy, world congress landmark vote, korea's ua, etc.) just not anything that upgrades specific improvements (i.e. economics which only improves custom houses and not other GP improvements). Additionally, the heads don't remove jungle when constructed on top of them, allowing you to also keep the jungle's science university bonus (and culture from the sacred path belief).