After the skappbeetles have escaped, make your way back down from Silas. Coming down from the war room, take a left onto the battlements and avoid the guard to find the skapp scurrying about on the floor.

From the ramparts, jump over the side of the battlements onto the tiled roof and head through the door. Make your way along the beams of the roof all the way to the far end to grab the skappbeetle.

Alternatively, if you've already lowered the ladder before leading to the beam, you can just climb that and the skappbeetle is right there.

This skapp is on the roof directly above Rolo. Jump to the roof from the wooden beam that you push to form a platform in the room with the collapsed roof.

This skappie is in a hidden area at the top of the room with the collapsed roof. Make your way into the fort above where Rolo is, and into the room with the collapsed roof.

Push the wooden beam through the wall and use the stool to climb through the window and outside onto the beam. Jump up the wooden platforms, and make your way back inside and onto the platforms there. On the wall to your left as you come in through the window is what looks like a window covered by some bars.

Use a stick or a bottle (there's a handy stick near where the rose is hidden up here) and throw it at the bars to make them swing down and form a ladder. Cliimb up the ladder and through the window to find the skappbeetle.

This is the same location as the chest with the ranger boots.

Across the courtyard from Rolo, head through the wooden door to the right of the long set of steps and pick the little fella off the floor.

In the east courtyard (across from Gusto and Fatale's cell), head up the long set of steps. The skappbeetle scurries around the corridor there.

This one is just scurrying about on the floor of the stables that are directly next to the main gate.

Note - it appears that the skapp may have been moved to the loft area of the stable. If it's not on the floor, climb the ladder and check up there (spotted by Mithraninja).